题目解析:
用occupied去进行定位,定位到该段最后一句:While this contest over an unoccupied nest site produced no obvious mortality, fights over occupied nests sometimes kill over 1,000 bees in a single battle.
从这句话可以看出,争夺空巢的战斗没有明显的伤亡, 然而争夺已占巢穴会造成尸横遍野。这和A选项匹配。BCD在原文中都没有提及。