The daytime quality of light in forests varies with the density of the vegetation, the angle of the Sun, and the amount of cloud in the sky. Both animals and plants have different appearances in these various lighting conditions. A color or pattern that is relatively indistinct in one kind of light may be quite conspicuous in another.Less colorful birds and animals that inhabit the rain forest tend to rely on forms of signaling other than the visual, particularly over long distances.
The piercing cries of the rhinoceros hornbill characterize the Southeast Asian rain forest, as do the unmistakable calls of the gibbons. In densely wooded environments,sound is the best means of communication over distance because in comparison with light, it travels with little impediment from trees and other vegetation. In forests, visual signals can be seen only at short distances, where they are not obstructed by trees. The male riflebird exploits both of these modes of signaling simultaneously in his courtship display. The sounds made as each wing is opened carry extremely well over distance and advertise his presence widely. The ritualized visual display communicates in close quarters when a female has approached.
